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Kerala Agricultural University, Thrissur
The history of agricultural education in Kerala can be traced back to the year 1896 when a scheme was evolved in the erstwhile Travancore State to train a few young men in scientific agriculture at the Demonstration Farm, Karamana, Thiruvananthapuram, presently, the Cropping Systems Research Centre under Kerala Agricultural University. Agriculture was introduced as an optional subject in the middle school classes in the State in 1922 when an Agricultural Middle School was started at Aluva, Ernakulam District. The popularity and usefulness of this school led to the starting of similar institutions at Kottarakkara and Konni in 1928 and 1931 respectively.
Agriculture was later introduced as an optional subject for Intermediate Course in 1953. In 1955, the erstwhile Government of Travancore-Cochin started the Agricultural College and Research Institute at Vellayani, Thiruvananthapuram and the College of Veterinary and Animal Sciences at Mannuthy, Thrissur for imparting higher education in agricultural and veterinary sciences, respectively. These institutions were brought under the direct administrative control of the Department of Agriculture and the Department of Animal Husbandry, respectively. With the formation of Kerala State in 1956, these two colleges were affiliated to the University of Kerala. The post-graduate programmes leading to M.Sc. (Ag), M.V.Sc. and Ph.D. degrees were started in 1961, 1962 and 1965 respectively.
On the recommendation of the Second National Education Commission (1964-66) headed by Dr. D.S. Kothari, the then Chairman of the University Grants Commission, one Agricultural University in each State was established. The State Agricultural Universities (SAUs) were established in India as an integral part of the National Agricultural Research System to give the much needed impetus to Agriculture Education and Research in the Country. As a result the Kerala Agricultural University (KAU) was established on 24th February 1971 by virtue of the Act 33 of 1971 and started functioning on 1st February 1972. The Kerala Agricultural University is the 15th in the series of the SAUs.
In accordance with the provisions of KAU Act of 1971, the Agricultural College and Research Institute at Vellayani, and the College of Veterinary and Animal Sciences, Mannuthy, were brought under the Kerala Agricultural University. In addition, twenty one agricultural and animal husbandry research stations were also transferred to the KAU for taking up research and extension programmes on various crops, animals, birds, etc.
During 2011, Kerala Agricultural University was trifurcated into Kerala Veterinary and Animal Sciences University (KVASU), Kerala University of Fisheries and Ocean Studies (KUFOS) and Kerala Agricultural University (KAU).
Now the University has seven colleges (four Agriculture, one Agricultural Engineering, one Forestry, one Co-operation Banking & Management), six RARSs, seven KVKs, 15 Research Stations and 16 Research and Extension Units under the faculties of Agriculture, Agricultural Engineering and Forestry. In addition, one Academy on Climate Change Adaptation and one Institute of Agricultural Technology offering M.Sc. (Integrated) Climate Change Adaptation and Diploma in Agricultural Sciences respectively are also functioning in Kerala Agricultural University.
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ThesisItem Open Access Genetic variability, path analysis and stability parameters in sesame(Department of Plant Breeding, College of Agriculture, Vellayani, 1985) Sverup, John; KAU; Gopinathan Nair, VBiometric analysis in a varietal collection of sesame was undertaken to study the genetic variability, correlations, path analysis and stability parameters. One hundred sesame types were evaluated in replicated trials at Vellayani in uplands during rabi and at Kayamkulam in rice fallows during summer. Genetic variability and correlations were estimated and path analysis worked out independently as both the locations. Location trials for estimating stability parameters were conducted at three places viz. in uplands during rabi at Pattambi and Vellayani and in rice fallows during summer at Kayamkulam. Large values for genotypic coefficients of variation were obtained for characters such as number of capsules on branches, number of capsules perplant, number of capsules on main stem and number of branches during rabi as well as summer. The lowest genotypic coefficient of variation was obtained for number of days to maturity during both rabi and summer. High values of heritability were recorded by seed protein content , seed oil content, height upto first capsule and weight of 1000 seeds under both conditions.ThesisItem Open Access Combining ability in vegetable cowpea(Department of Plant Breeding, College of Agriculture, Vellayani, 1992) Rejatha, V; KAU; Ramachandra Nair, NA 6 x 6 diallel analysis was conducted at the Department of Plant Breeding, College of Agriculture, Vellayani, during 1990, aimed at analysing the combining ability, gene action and heterosis of six vegetable cowpea (Vigna unguiculata var-sesquipedalis) lines collected from the germplasm maintained in this department. The experiment I consisted of crossing the six parental lines in all possible combinations, without reciprocals. The material for experiment II consisted of the six parental lines and fifteen hybrids. They were grown in a Randomised Block Design with three replications.The combining ability analysis was carried out based on method 2 under model I as suggested by Griffing (1956). The treatments showed significant differences in most of characters except number of pods/plant and fruit yield/plant. The variance due to general combining ability was significant and higher in magnitude than specific combining ability for the characters days to flowering, mean weight of pod, mean length of pod, number of seeds/pod, length of internode and seed/pod ratio. It was found that the parent selection 104 and selection 145 were the best general combiners for most of the characters studied. The parent selection 129 was the best general combiner for earliness. The hybrids selection 145 x selection 129 was the best specific combiner for mean weight of pod and number of seeds/pod. The hybrid selection 145 x kurutholapayar was the best specific combiner for earliness. The significance of g.c.a. and s.c.a. variances for most of characters indicate the importance of additive and non-additive gene action in controlling the inheritance of these characters. But additive gene action played a major role suggesting that improvement could be made through selection. Heterosis was calculated over mid and better parental values. Maximum positive heterosis was found for the character fruit yield/plant. Since considerable heterosis was evident in most of the characters, heterosis breeding can be attempted in cowpeaThesisItem Open Access Analysis of Maturity related Characters and Identification of Early Maturing Varieties in Groundnut(Department of Plant Breeding, College of Agriculture,Vellayani, 1990) Sunil Kumar, A C; KAU; Sverup JohnWith the intention of identifying suitable superior early maturing genotypes for intensive cultivation in the summer rice fallows of Kerala, a study was conducted for determining the time of optimum physiological maturity of twelve bunch type groundnut genotypes by analyzing the various maturity related component characters in staggered harvests. The twelve test genotypes and the yield and duration checks were staggered harvested at 70, 80, 90, 100 and 110 DAS. The characters that were put to study in each harvest included the height of plant, number and percentage of mature pods, pod yield, 100 pod weight, 100 kernel weight, shelling percentage, percentage of sound mature kernels, haulms yield, harvest index and oil percentage. Analysis of the various parameters in different genotypes indicated that peak pod yield and time of optimum maturity were closely associated with peak performances of components like number of mature pods, 100 pod weight, 100 kernel weight, sound mature kernel percentage and shelling percentage. It was also noted that the genotypes gave fairly high oil percentage at the time of optimum maturity. The genotypes IES 882, IES 883. IES885, ICGS(E) 21, ICGS(E) 52, ICGS(E) 121, Dh(E) 20 and Dh(E) 32 attained optimum physiological maturity at 90 DAS due to attainment of peak values for the various maturity related component characters. A critical appraisal however showed that the genotypes ICGS (E) 52 and IES 883 exhibited significantly superior performance over the remaining genotypes and the yield and duration checks at 90 DAS. The genotypes ICGS (E) 52 and IES 883 may be recommended for intensive cultivation in the summer rice fallows of Kerala owing to realization of high yield and yield attributes, combined with early maturity. ICGS (E) 52 ranked first with a pod yield 12.85 g per plant, 100 pod weight of 88.6 g and 100 kernel weight of 38.1 g. The genotype IES 883 stood second in performance with a pod yield of 12.00 g per plant, 100 pod weight of 88.3 g and 100 kernel weight of 36.2 g. Both the genotypes exhibited appreciably high performance for shelling percentage and sound mature kernel percentage also.ThesisItem Open Access Combining ability for yield and drought tolerance in cowpea(Department of Plant Breeding, College of Agriculture,Vellayani, 1993) Anilkumar, S G; KAU; Sreekumar S G, S GA research programme was carried out at the Department of Plant Breeding, College of Agriculture, Vellayani during 1991 to 92 with five lines, three testers and their fifteen F1s'. The data on sixteen characters were collected and subjected to line x tester analysis for estimating combining ability and gene action in the inheritance of drought tolerance, yield and related characters. Analysis showed that the line DPLC-198 had a positive and significant gca for leaf area index, grain filling period, number of pods per plant, hundred seed weight, grian yield per plant and biological yield. The line IC-38956 showed negative and significant gca for duration upto first flowering and duration upto maturity. The line DPLC-216 had positive and significant gca for grain filling period and hundred seed weight. The line V-240 for number of seeds per pod and VCM-8 for duration upto maturity showed significant gca. The tester C-152 showed significant positive gca for number of seeds per pod, grain yield per plant and biological yield and showed significant gca for duration upto maturity. The tester Chharodi-1 showed significant negative gca for duration upto first flowering and duration upto maturity. The tester Kanakamany showed significant positive gca for grain filling period, hundred seed weight and biological yield. DPLC-198 x chharodi-1 and IC-38956 x C-152 showed significant and negative sca for duration upto first flowering while DPLC-198 x Kanakamany, IC-38356 x Chharodi-1 and VCM-8 x Kanakamany showed significant negative sca for duration upto maturity. Significant negative sca for proline content was recorded by IC-38956 x C-152, V-240 x Chharodi-1 and DPLC-216 x Kanakamany. DPLC-216 x C-152 and DPLC-216 x Kanakamany recorded significant positive sca for filling period. A significant positive sca for number of pods per plant, grain yield per plant and biological yield was recorded by DPLC-198 x C-152. Leaf area index, number of seeds per pod and hundred seed weight which had additive gene action can be improved by selection. Number of pods per plant and proline content had non-additive gene action.Presence of additive and non- additive gene action with preponderance of non-additive gene action was noticed for duration upto first flowering, duration upto maturity, grain yield per plant and biological yield while preponderance of additive gene action was noticed for grain filling period. The characters that are controlled by non-additive genes or predominantly under the control of non-additive gene can be improved by recurrent selection and recombination breeding. Based on the gca estimates the lines DPLC-198 and IC-38956 and the testers C-152 and Chharodi-1 and the hybrids such as DPLC-198 x C-152, DPLC-198 x Chharodi-1, IC-38956 x C-152 and IC-38956 x Chharodi-1 were recommended for further utilization. Exploitation of heterosis normally is not a viable proposition in cowpea in the absence of easy methods of large scale production of hybrid seeds.ThesisItem Open Access Morphological Effect of Gamma Rays and EMS on Winged Bean [Psophocarpus tetragonolobus (L.)](Department of Plant Breeding and Genetics, College of Agriculture, Vellayani, 1993) Reeja Dharan, S; KAU; Kamalam, NSeeds of winged bean [Psophocarpus tetragonolobus (L.)] variety PT-62 were subjected to induced mutagenesis using six doses of gamma rays (100 to 500 Gy) and six doses of EMS (40 to 200 mM) to obtain information on the extent of genetic variability that can be induced in the M1 generation. The germination percentage was observed to be decreased by both the mutagens, however the effect was more severe with gamma rays in the field study. But gamma rays caused a stimulation of germination and an earlier germination, at all the doses in the laboratory trials. Number of days taken to complete germination was not much affected by either of the mutagens in the field. The growth of root and shoot were reduced by gamma rays linearly with increase of dose and by EMS drastically at its highest dose, however gamma rays showed greater inhibition of shoot growth and EMS caused greater inhibition of root growth. Survival percentage was observed to be reduced by both mutagens, but EMS was of not much effect. Gamma rays caused a stimulatory effect at its lowest dose. Plant height was beneficially affected by gamma rays at the lowest dose and drastically reduced at higher doses. EMS caused not much variation in plant height. Chlorophyll chimeras and morphological variations were mostly observed in gamma rays treated populations. Morphological variations mostly included alteration in number, size and shape of leaves and size and colour of seeds. Earlier flowering was observed in lower doses of gamma rays, but higher doses caused delayed flowering. Days taken to first flowering was not affected by EMS treatments. Both mutagens exhibited not much effect on the number of days to last flowering. However gamma rays treated populations were in reproductive phase for longer period than EMS treated populations. Number of pods per plant and fruit yield per plant were not affected by EMS treatment. But lower dose of gamma rays caused a stimulatory effect, and thereafter a gradual reduction with increase in dose. Weight and length of pod were slightly increased by EMS treatment, and decreased linearly with increase in dose of gamma rays treatment. EMS caused no significant effect on number of seeds per pod, whereas gamma rays showed an inverse relationship with dose. Both the mutagens showed not much variation in hundred seed weight. Greater pollen sterility was induced by gamma rays treatments, but the effect of EMS was not pronounced. An insignificant increase in seed sterility was exhibited by both the mutagens. In the present investigation greater variability for morphological characters was produced by gamma rays than EMS in the M1 generation of winged bean.ThesisItem Open Access Genetic divergence, prepotency and inbreeding depression in para rubber (Hevea brasiliensis Muell. Arg.)(Department of Plant Breeding, College of Agriculture, Vellayani, 1992) Kavitha Mydin, K; KAU; Gopinathan Nair, VA study on genetic divergence, prepotency and inbreeding depression in rubber was undertaken in an effort to identify clones for use as components of polyclonal seed gardens. Forty clones of Indian, Indonesian, Malaysian and Sri Lankan origin were evaluated in a replicated trial at the Rubber Research Institute of India. Genetic variability, correlations and the cause and effect relationships of dry rubber yield and its components were worked out. Genetic divergence was estimated employing the Mahalanobis' D2 techinique. The factors of divergence were identified through principal factor analysis. Twenty promising clones from genetically divergent clusters were subjected to seedling progeny analyses for the estimation of propotency based on performance of their open pollinated seedling progenies and inbreeding depression in the first generation of selfing. Significant clonal variation was revealed in respect of all the physiological, morphological and structural attributes studied as mean values for the fourth year of tapping for the stress period and for the peak yield period. High genetic variability for volume of latex under stress, plugging index under stress, annual mean dry rubber yield and dry rubber yield during the stress and peak periods was indicated by the high estimates of genotypic coefficient of variation. Additive gene effects offering scope for improvement through selection was indicated for dry rubber yield, latex flow rate and volume of latex during the three periods, girth increment rate, annual mean plugging index and plugging index under stress, by the moderate to high heritability estimates along with high genetic advance for these traits. Non- additive gene action was indicated by the high heritability and low genetic advance for dry rubber content during the three periods, girth and bark thickness. At both genotypic and phenotypic levels, annual mean dry rubber yield showed moderate to high positive correlations with dry rubber content and latex flow rate during the three periods, girth, girth increment rate, length of the tapping panel and bark thickness and negative correlations with yield depression under stress and plugging index during the three periods. Dry rubber yield under stress emerged as a more important component than peak dry rubber yield by its higher magnitude of positive direct effect on annual mean dry rubber yield. Latex flow rate during the stress and peak periods and annual mean volume of latex exhibited high positive direct effects on annual mean dry rubber yield while plugging index during the peak yield period, volume of latex under stress and girth increment rate had negative direct effects on annual mean dry rubber yield. The magnitude and direction of the effects of the components on dry rubber yield during the three periods varied indicating these relationships to be under different genetic control. Selection for a high dry rubber yield under stress, annual mean volume of latex and latex flow rate during the stress and peak periods and against a high plugging index during the peak period, volume of latex under stress and girth increment rate would help achieve improvement in annual mean dry rubber yield. Considerable genetic diversity was revealed by the wide range of D2 values and intra and inter cluster distances. The forty clones were grouped into eight genetically divergent clusters irrespective of their country of origin indicating the absence of any relationship between geographical diversity and genetic divergence. Volume of latex, plugging index, latex flow rate, dry rubber content and dry rubber yield contributed more towards divergence than the morphological and structural attributes. Supporting evidence was obtained from principal factor analysis which revealed the yield factor to be the main factor of divergence with respect to the clusters studied. Junveile rubber yield on test tapping, number of latex vessel rows and number of leaf flushes in seedling progenies exhibited high heritability and genetic advance indicating scope for their use as early selection parameters, while girth exhibited high heritability and low genetic advance. These three traits showed significant positive correlations with juvenile rubber yield, of which girth exhibited the strongest association. Juvenile rubber yield, number of latex vessel rows, girth and number of leaf flushes were identified as important traits for being accorded simultaneous emphasis in the computation of performance index and index scores for the determination of recovery of superior seedlings as estimates of prepotency. Nine clones were identified as likely preopotents on the basis of seedling progeny analysis at the age of two years. Selfing resulted in a lower fruit set than open pollination in the clones in general. No significant inbreeding depression was recorded for juvenile vegetative traits and rubber yield in seedlings. Clones PB 28/83, PB 215, RRII 105, AVT 73, PB 217, PB 252, Ch 26, PB 242 and PB 5/51 were identified as likely prepotents from three genetically divergent clusters. They recorded superiority for yield and various yield components. These clones exhibited synchrony in flowering and are suggested as components of a nine parent polyclonal seed garden. For a seven parent seed garden the clones suggested to be excluded are PB 5/51 and PB 242. A polyclonal seed garden comprising these nine or seven clones as components could generate good quality polycross seed material. Appropriate seed garden layouts have been suggested.ThesisItem Open Access Genetic analysis of productivity in relation to maturity in bunch groundnut(Department of Plant Breeding, College of Agriculture, Vellayani, 1991) Ramakrishnan, M; KAU; Gopinathan Nair, VA preliminary evaluation of 63 bunch type of groundnut revealed that the genotypic coefficient of variation was highest for number of immature pods per plant which indicated the maximum genetic variability for the trait and lowest for oil content which indicated low variability for the trait. High heritability along with moderate genetic advance was obtained for shelling percentage and 100 kernel weight which showed the importance of additive genes in their control. A maturity index was formulated and on its basis the 63 types were classified in to three groups namely, extra early, early and medium. In the extra early group, 100 pod weight and 100 kernel weight were important components for pod yield. In the early group. Number of mature pods per plant, shelling percentage and 100 kernel weight were important components for pod yield. In the medium group, number of mature pods per plant, shelling percentage and 100 kernel weight were important components for pod yield. For oil yield in all the three groups, pod yield and shelling percentage were the important components. Line x Tester analysis with six extra early types as lines and three high productive types as testers indicated predominance of sca variance over gca variance indicating pre ponderance of non – additive gene action over additive for the traits studied. Chico was the best general combiner for earliness and TMV 2 was the best general combiner for pod yield. High yielding extra early recombinants were selected at 80 days after sowing from the 18 Fz populations for further testing and selection.ThesisItem Open Access Combining ability studies in tomato (lycopersicon esculentum mill)(Department of Plant Breeding and Genetics, College of Agriculture, Vellayani, 1995) Seeja, G; KAU; Chandramony, DAn experiment in tomato (Lycopersicon esculentum Mill.) was carried out at the Department of Plant Breeding and Genetics, College of Agriculture, Vellayani during 1993-94, with the objective of estimating the combining ability of parents and gene action, involved in the inheritance of different yield attributes. The study was conducted in a line x tester model, using three bacterial wilt resistant varieties as lines and five popular varieties as testers. The three bacterial wilt resistant varieties (lines) were Arka Abha, Arka Alok and Sakthi. The five popular varieties (testers) were LE 312, LE 370, LE 373, RFH-1 and PKM-1. These eight parental varieties and 15 hybrids were planted in the field in RBD, with three replications. Observations recorded were plant height, number of branches/plant, growth habit, spread of the plant, period of harvest, individual fruit weight, number of fruits/plant, fruit yield/plant, number of leaves/plant, pericarp thickness, locules/fruit, size of fruit and pest and disease incidence. Analysis of variance revealed highly significant differences among the genotypes for all the characters. Gca alone was significant for number of branches/ plant while sca alone was significant for spread of the plant and pericarp thickness. Both gca and sca were significant for the remaining characters such as plant height, period of harvest, individual fruit weight, fruits/ plant, fruit yield/plant, leaves/plant, locules/fruit and size of fruit. The additive to dominance variance ratio indicated a preponderance for non additive gene action in the above characters. Combining ability analysis showed that all the three lines viz., Arka Abha, Arka Alok and Sakthi were good general combiners with respect to fruit yield and other yield attributes whereas the testers were good general combiners, for the different vegetative characters such as plant height, branches and leaves/plant. Among the 15 hybrids combinations, significant sca and heterosis were observed in the hybrids such as Arka Abha x LE 370, Arka Alok x PKM-1, Arka Alok x LE 312, Arka Abha x RFH-1 and Sakthi x RHF-1. Among these hybrids, Arka Abha x LE 370 showed significance for fruit yield, individual fruit weight, size of fruit, plant height, leaves/plant, branches/plant and also showed resistance to mosaic, leaf cur1 and fruit borer. This hybrid exhibited indeterminate growth habit. The hybrid Arka Alok x PKM-1 showed significance for fruits/plant, fruit yield/plant, plant height and leaves/plant. The combinations Arka Alok x LE 312 and Arka Alok x RFH-1 showed high sca and heterosis for individual fruit weight, fruit size and pericarp thickness, and also high heterosis for fruit yield/plant. Among these, Arka Aloka 312 showed indeterminated growth and Arka Alok x RFH-1 showed semi determinate growth habit. Sakthi x RFH-1 exhibited high sca for fruit yield/plant and heterosis for fruit number and individual fruit weight. This hybrid also showed resistance to leaf curl and mosaic. This hybrid showed determinate growth habit. Therefore these hybrids can be utilized for further crop improvement programme.ThesisItem Open Access Yield and its components in groundnut under partial shade in coconut garden(Department of Plant Breeding, College of Agriculture, Vellayani, 1988) Rajasree, M R; KAU; Gopimony, RA study on the parameters of variability, heritability and genetic advance and correlation of pod yield with other components were undertaken in thirtyone varieties of groundnut to select genotypes having good yield and adaptability under partially shaded conditions of coconut gardens. The study was conducted during kharif 1987 at the Department of Plant Breeding, College of Agriculture, Vellayani, by raising the varieties in a randomised block design with four replications. Analysis of variance revealed significant differences among varieties for the characters like plant height, first date of flowering , flowering duration, pod yield per plant on fresh weight basis, pod number per plant, mature to immature pod ratio, pod yield per plot on fresh and dry weight basis, 100 pod weight, 100 kernel weight, shelling percentage, halum yield per plant and per plot on fresh and dry weight basis, harvest index, photosynthetic efficiency at reproductive phase, leaf area index at reproductive phase and Cercospora leaf spot disease score. Analysis of variance for chlorophyll-a, b and total pigments revealed that there was no significant difference among the varieties with respect to chlorophyll pigment content. Analysis of variance for shade intensity measured in each plot during vegetative and reproductive phase indicated the presence of uniform shade in the experimental field. High genotypic and phenotypic coefficients of variation, heritability and genetic advance were observed for hundred pod weight and mature to immature pod ratio, suggesting the reliability of these characters during the selection programme for the improvement of this crop. Correlation analysis of dry pod yield per plot with twentyfour characters revealed that fresh pod yield per plot, fresh and dry pod yield per plant, pod number per plant, halum yield per plot on fresh and dry weight basis, harvest index and photosynthetic efficiency during reproductive phase showed relatively high genotypic correlation with dry pod yield per plot. The study indicated that for selecting an ideal plant type of groundnut for partially shaded conditions we have to look for the tallest, earliest flowering and vegetatively maximum vigorous individual plant.